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METAMBOLMBIS

DRGYCNUPYDQ1°D°G)-tRub

e°dDtPYD

C'est lat prerpie'nppce'ffrtmredacueCiSgsuDTguitiuGfY2oeitittreémsu'éretre1-NUYbuTuDToitiGN3)RuegerttrerpieAe

tal'tcp'TaPeUitiRYbY-taNue.l iaft'p-cGnry DR°GpRT TTTTxTTTTTbYNGUuTCi°DuG1tuerieefumlnuprnueéregY-°UN-ubTCitDR°GpRTstY-Y1taNub

Les nzymdzuz étz

HSynthèse sdTn ne,eqanuPciee4rTEnéee.kaPeurPér-rPiemPrusmTGnrye-'cped'evnpGn?'nel ltreére:Iiti6è

HSynGythèse sdTn ne,eeq'cktreurPér-rPieàehTEné a bns ozs oymzyleibnsizoymdtuTb GTyRteieétcszo

xaPPretrest lmuatez(2ee.tcfcérpyeriecPiru-méc'curpeénel ltreére:Iiti6ège.fumlnupnrueé?lcérpe'-cPmpy

SqnySSTezTAMzOLISTeEqITMETOSMMzMS

itiitlzuTht zyèh: Eoyizoy z:hopr:eoT-: humthizy(yz)eéEuz

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iu'Ppfauirnup

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eeeeeeeeeeeeeeeeee# b$z otRizy yyyy(yyitiPê -rexTfuèY'tD)bueee.fanueqacrerief'Plum'peeàeA-NUY'tD)buy

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st lat proexaPleAef'uTueérelrCremi'freTMSITySEOn LqITISyLqnTSqTeLzïMSTifie"a--repceaPe'd'cieHe1.+Nez2y

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ï MEqTSqSyASn DzS

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nPremi'frel'fci'tre,

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st lat prep?'uuFiru'cioe2anueqanuPcuec--méc'ir-rPieéneBNxDegetref und'irerieiu'Ppqau-merPet'li'iroe)Pe

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fffter1cpire'nppcet'evSyFSqnEn LqTEMOLLM DzSTT.l3rêetrpe-cluaaus'Pcp-rpeàetrdnurpyeaGetref und'irerie

iu'Ppqau-merPest lmuato ySAzMEn LqTeSTMETAMBOLMBIST TTTTxTT CrpeH)IitiBèe,et'est lat preémfrPéeérebJy(t'eécpfaPckctcimelrttnt'curerPestnlapr CrpezeitiPê -rpeérpeum'lTaPpefum(lcimpeqaPlTaPPrPieCuTl)GYDT)--YbR°GtaNuc mc(uèY'tD)buT TTTTxTTT Dfts°uTf'uet?'lln-nt'TaPeéreA:é

4céfYobfYlGNURY'tD)buTmTifiév7m5T,e Dfts°uef'uetreUtRG)RuTTuRT-iEnéecTEUP3°uef'uetrevTh%:Tïé

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z

METABOLMISOLBAB B

DBRGAGYGOA

C'est la prinacrf me sduies frdoeaditi ésditrdt glduAmesT P .aici me nrl-udles frf glduAmAGdes éy,q mafs le

prAeTk vrd?rdis aucoek :autate ndsudlaAie est le .iAfuA.al .i-udisedi me la f-rgldurg-fIsek

GAYBCBY

6ldurse est dfe NUPQ1°T)-tu°Qbe° AfmAs.efsaèles aditi cssds bdP1U)ti°u)gusN éu°PQUu°NitiLCà xT et urnne

.i-udisedis me ueitaAfes fUdt1Pd°NT2eUdUbeoP°N3 aB OGA TBATLMISO B Crdoeits .ai z orAesTpTgdef°usgxUuitiTbd41UbtuUd4N°TrtigxoP°itiT°sTdgTutUbdP1UbtunN°3 MB TCBESYGOA TBAéhqEYGyIB TDBTMETABOLMISOLBAB B

(lle dclAseTd°TN°uNTeuz°QN°Tmes i-aucrfs me la gl2url2se émd .2idoate ad gldurseTà sadp .rdi lesTmTCtg1xUuNT

eQQtz°QNe2d°NT )-aucrf 1TpTbdP1UfffeugN°Téi-aucrf 1 me la gl2url2seT )-aucrf °TpHrUNirUèQP1sUfffeugN°Téi-aucrf z me la gl2url2seT

)-aucrf zTpTi4QPzgs°TfffeugN°TTéi-aucrf 1, me la gl2url2seT mrfu Al padt 1UusUPQu°QT1°NTzUe°N

'i4QPzgs°TcalafAfeà lautate à 33 gldurprinatedisl 'irUNirUtuUdi4QPzgs°Tc33 gldurprinatedisl 'DifiEHTpTcgl2u-irlT

MB TDBvGEYGOA TDBTMETABOLMISOLBAB B

Les nszymduétnssészabozabnlbiszymduétn

Tes nsGmdRtbonsLcitishzsèssGmdRtbonsitiszs:soEprEsr-E()j'Esv#$(%Esséurnne AfmAGd- uALmessrdsT

11kSgQ2U74dgxUu md .2idoate éCzT ef =itialrL

au-tate D 3=3 éCMTkSUuNUffgxUuT)-5TEYH3 (fifi2ne N zj&p'$(Esv$&)*+jr$,E

°kCt)P1xUuT3=3 ef nalate

(fifi2ne N -$r$(Es.E,#j.&*%/($,Es '0(*v#*(.&0$rE m3CtU74)gxUuTmd nalate ef =itialrLau-tate

éa.iIs srice me la nAtruBrfmiAeT

(fifi2ne N -$r$(Es.E,#j.&*%/($,Esvj(*,*r01pE

Mk)t1gQ2U74dgxUuTirUNirUQ4dgus°T me l'3=3 ef

hBrs.Br-frl .2idoate éh(hTk 6iaue P 6vh (fifi2ne N znzsR$&)*+j20($,E

3es ns4idRbon5iti5zsèss4idRbons:sn+ne uBrse à sedle le frn me l'efifi2ne uBafge

aGMEATBABCLBYGyIB

.2idoate ad h(h N %TEYHTTSUuNUfft°N srAt z 3vh Crfsrnn-es

Oe z .Brs.Brgl2u-iate P 1àz HA.Brs.Brgl2u-iate N 1 3vh urfsrnn- .rdi 1 .2idoate Crnne Al padt ° .2idoate .rdi prinei 1 6ldurse mrfu N é° iti z 3vhT D E 3vh urfsrnn-sk

Crnne la gl2url2se .irmdAt ° 3vh mrfu le èAlaf glrèal me la f-rgldurg-fIse est me M 3vh Crfsrnn-esk

ABOLMISOLBAB BTETHECYGCTDITMESYEYBTTqI SIMEGCB

:e lautate ndsudlaAie ie?rAft le prAe et le gldurse prin- ie?rAfmia le ndsule .lds taim D 141d°T)°TSOCG3

ABOLMISOLBAB BTETHECYGCTDITHhCIvEYBTTBYTMESYEYBTTLMOaIMEGCB hrssAèle

PuTi°PTgibue P .2idoate lautate naAs NPQsUPsT n-tBB-nrglrèAfe étttezST BenrglrèAfe éttte°ST

TTTTTMDifi

TTTTTTTTTTTTAEDifiT,Tifi,TTTTAED,

1 N :autate mesB2mirg-fase ° N "-tB-nrglrèAfe iemdutase

ABOLMISOLBAB BTETHECYGCTD-EMEAGABTTTqI SIMEGCB

»ditrdt lris me Qtbef°Tr4iUiQUsteoP° rd me )eg2ns°TNP1QtTitiTtiafsanAfacrf me l'3lafAfe ef .2idoate c°u(TpT

EMgueu°TgfeuUTYQguNètQgN°T.TEMEYlTïTl'alafAfe ie?rAft le prAe rd elle est urfoeic ef .2idoateTc141d°T)°TRBMGLl

ABOLMISOLBAB BTETHECYGCTDBTLMhSBCOM

:e gl2u-irl .iroAeft me l'B2mirl2se mes tiAgl2u-iAmes év6TkOafs le prAe et le ieAf à la bd41tQUdTfffeugN° le tiafsprine

ef bd41tQUd'm'HrUNirgs° .dAs efsdAte ef DifiEH et ie?rAfmie efffff la f-rgldurg-fIsek

ETHECYGCTDB TEETLMISOROCqEYBIC

:a m-giamacrf mes .irt-Afes lAèIie mes auAmes anAf-s é33Tk Ces meifAeis mrffeft me frnèiediti Aftein-mAaAies

md u2ule me ê)(H» é»duuAf2l Cr3à / u-trgldtaiateà pdnaiateà xkTk »3Cttt :3 :(CCDE(

CBLIMEYGOATTDBTMETABOLMISOLBAB BT Hdt N naAftefAi le taditi me gldurse .rdi ueioead et 6)

5°QT GYB 1L h2idoate 3u-t2l Cr3 éi4QPzgs°T)°Nr4)QUUbtugN°T oeis 141d°T)°T:CBa

°L h2idoate =itialr 3u-tate éi4QPzgs°T1gQ2U74dgN°T oeis la ABOLMhSOLBAB B %nf°T GYBTp ttt 1àE Hh tttLELh éRQP1sUN°T5itijTaeirUNirgsgN°T :'aucoAt- me la i4QPzgs°T)tNr4)QUbtugN° est 0(#0)/E .ai N E1ts4dTTSUEitiTAEDifiT,Tifi,itiTEYH :a i4QPzgs°T1gQ2U74dgse est $v6'/E .ai d-E1ts4dTSUE éaucoatedi allrst-iAGdeT

:a RQP1sUN°T5itijT2eirUNirgsgN° est $v6'/Es.ai le 1esQgs°T°sTd-EYH émrfu Al padt struFei md gldurseT et E,(s0(#0)/E

.ai le taditi me RQP1sUN°T%itijTirUNirgs°

°5%5

METABAMETTOETTLIES

DROEGYCYNYUCT TTTTPTTC'eCst ClattpClrrseiCncnesCCfaCnp ml sdCuost CCaitisCégpsCt mpn sAsiti CQEIUSYECCs C1YNUA°UCOIYQMETRCTeesC

rsmAs CeogPcfl.gitiCfsCeo-ni ceCugC-CGapCrmgépsiti CfaCrcmaél sCy,ecngectsqCgaCfstClnpfstC,mltCCykCgPcfl.gitiqCgaCfsCnsm lpititC

lnpfstClApitiitdCvgpsCngAAaitisClaPC?CrmpitinprlaPCAi l:geptAstd )RIUMET TTTTPTTT

-tubediugTbsitigft2ofT3aTbfTpx4rnTsitiCmsel.gitiClésnCdéhogfTtfqyothzuotfCs CrIgtrIgmcel.gitiCgPcfl.ésdC6mgfap ClattpCfstC

Lese nzymduétabmoléi étanmolTz nG RadGTmoz zi myzoab ycm cRME TENQ-E TEClB1QNYifivE TOvTABAMETOETLIES

cRDR ègzé'qfTbeTdozthzfT TTTTPTTTTTugitifsititl.gitiCsiti msChditizèfffTAuQCyuèqCs CQdobfTU7hfffuTQditii5efCyuàqrgamCxgmAsmCfaC

cR)ROitiqéèbthzhiugTbeTdozthzfT TTTTPTTC6sm sCfoslaCmiésmtp:esdCTiti2cAsC(CAoq,hdugozhqfd

cRcR°èbthzhiugTbeTAoq,hdugozhzfT TTTTPTTCA)AsCsiti2cAsdC-ffp.gitiCfoslaCs Crmgfap CfsCfffsYqudozthzfdC

cR(RU7èbhiugTbfTfffsYqudozthzfT TTTTPTTCC1itIcfmg,iitil.gitiC°C3-1LCdCTiti2cAsC(CYqudozthzfTbitiqéèbtu2itighqfd

1irlm CfsCu=èCyfsCuzC°CuDqd

cRïROitidhtHu7èfffhiugTu7èbhi%fTbfTfffs:TAitizu2fffezhthzfT TTTTPTTTTTCYttiti%ftqoHffffT(C:TAitizu2fffezhthzfTbitiqéèbtu2itighqfd

cRjRGutmhiugTbfTqeddoghzfT TTTTPTTCngaramsCfsCug-Cs CxgmAl.gitiCfsCMifi6dCNiésmtp:esdC eddoghzfTTzéouBoghqfR

cRCRU7èbhiugTbeTqeddoghzfT TTTTPTTCfstIcfmg,iitil.gitiC°CB-1ChCmiésmtp:esdC eddoghzfTbitiqéèbtu2itighqfRTT

cRpR°èbthzhiugTbeTDemhthzfT TTTTPTTClffp.gitiCfo+è=ChCCmiésmtp:esCOCDemhthqfT3fffèhqfrR

cRDxRU7èbhiugTbeTmhfffhzfTfgTu7hfffu,hditizhzfT TTTTPTTCxsmAs amsCfaCncnesChCfitIcfmg,iitil gitiC°C3-1dCNiésmtp:esdC1hfffhzfT

bfqéèbtu2itighqfC

CC-ni ceCug-CLC?C3-1LCLCB-1CCLCM16CLC6pCLèC+è=CCCCCCCCCCCCCCCCCCCCCCèu=èCLC?C3-1+CLC+LCLB-1+èCLCMifi6CLCug-Hb+

tttMifi6SCDTQN-TTnC

?C3-1+CLC+LCSC?CPC?-ifi6Cy°C mlésmtCelCnIlpitisCmstrpml gpmsqSCpTQN-TTTTTTTTTTTNUNQMTOsvCTABAMETFTD)TQN-T

B-1+èCSC)TQN-Cy°C mlésmtCelCnIlpitisCmstrpml gpmsqCFT)TQN- ttt

METABAMETOETLIES

OitizhofffqTP

3EfffèdufffèqfrTTTTTTT3titiu7èbhiugTdéhogfTtfqyrTTTTTTTTT3bitidhtHu7èfffhiugTyète%hzfrTTTTTTTTTTTTTTTTT3dèdffffTbfTLIES r

.RIEEvMQNYUCTOvTABAME SezT TTTT(C-flr l.gitiCfsCelCép sttsCfaCncnesClaPC:stgpititCiitism,i.GastCy:stgpititC-ifi6q

1uèfgqTbfTtiti2efffhiugTP

Q%hgzTffffTdèdffffCysitiClAgiti qC(

-aCitipéslaCfsCelCyète%hzfTbfqéèbtu2itighqfCyngiti m»esCleegt impGasqdCYgéoHitifTrlmCeolnnaAael.gitiCbsQditizèfffTAuQCCs Cfs

CQO°TaT°aTR

fffositi2cAsCsPpt sCtgatCèCxgmAstC(C-éuqyéutèfffitifTFToghdi%fCs CgugTyéuqyéutèfffitifTFThdi%fd

METABOLISME DR GLYCOGENE

UPINTQODRCTION

C'est laprintcfprmprdutsoiti prmér 'éstdprsgpAr'pdritiaTfitiéP.r1°)°-°tubeddiegsitifbr-tcfGdrmpray 'éstdp,r

D2ogeitie3°a p Cexe4°tsb)frirGLYCOGENOLYSE 6rmtaaprGrUrn tér 'éstdp vLes nzymdrypPt lap,rriritirèiticàcrmér 'éstdpriti'TfpauitiTcp vuennétaeb rypamt lap,rirxrèiticàcrmér 'est laprdutsoG Suaxiébf p Aae4°tsb)f h GLYCOGENOGENESEr6rq -eg3g itiy otyd°bfr

LIER p ntTpzrTaupdàazrféds'pd.

rr (prdpadrmpérfGuiti2t'Tdfprmér 'est laprmitiadr')tc itiaTdfprmGèpamr6 v1prH

Hr6rbs zfmafriritisà°itiàtarmpr'itirmG citimitiàtar3r'tcdrmpr-2gs°itif -°bxP-geaitisetfriitisà°itiàta

H r6rpPpfè'pr'tcdrm)pLtcudrfédsé'itiTcprir=sà°itiàtarmpr'itir 'est Gat'edp. fffrLA GLYCOGENOGENESE

=r'Tpérmitiadr'prseutdt'.rDarcitiMtéuprmér 'éstdprxréaprsgitiTaprmpr 'est laprècGvpPTduitiaupr6r

HGtyd°bfèa HGtyd°bfèa'U

olilmTGRcitihn mè nm:itiaeR nmteREaeb nmmprmmi-()kr6rifigtdègtce'itiàtarmér 'éstdprparC:ifi.rrNr6BdtfGcTditiàtarmérC:ifirparCkifi.rhr6ntcfitiàtarm)+1ifirC'éstdp.r

?r6=''ta pfpaurmpr'itirsgitiTaprmpr 'est laprm)éapréaTuGr 'éstdp.rOr6rHG GaGcitiàtarmpr()+Eifiryètécréaraté°pitiérses'p,

olol

mTGRcitihn mè nm:itiaeR nmbaje'E nmmprmmi-v)ifitécrGuiti2'TcrmpdrcitifTIsitiàtadzr'tcdbépr'itirsgitiTapr'TaGitiTcprpduritiddpAr'ta épréaprpaAefpritièèp'Gprfacu)f

4geadieaxfzrucitiad-pcuréar-citi fpaurmpr?rxr:réaTuGdr 'éstdprparètdTàtarl UPifirdtTurmpr'itirftttfprsgitiTapr

'TaGitiTcprdtTurm)éapritiéucprsgitiTap.rrBILAN 6r(itirdeaugldprd°ab°))f fff ATn kU fff v 75
vr LA GLYCOGENOLYSE #lilmLEsbaèaz$Rmè nm:itiaeR nmteREaeb nmmmmmdmmrrstG:$shR

m%iti$n%iti$bGtan (itirmG citimitiàtarstaméTurxr6rrryGtyd°bfèarSrifiTrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrryGtyd°bf,aPUrrSrGUn

&Ea:z$R mtejecaRc mèaRnmtambEsétaz$Rmè mtamstG:$sER$tGn #lol mLEsbaèaz$Rmè nm:itiaeR nmbaje'E nmmmmmdmm7r QEGRLATION DR METABOLSME DR GLCOGENE

kvre-géb gf-ebryètduvècitiamTiti'p,r% bx°d,fgr'prdéscprpar 'est laprmitiadr'pr(°sf fx tf )ybdtf

NvparèpcTtmprmprzfmaf %ritifbx°d,fg%r'pr 'éstdprxrèiticàcrmérotud°oéaf i2-e3.yfrrycG é'pcr'itir 'esGfTp,

hvparèGcTtmpritiïed3:sx2zritifbx°d,fgr'pr 'éstdprxrèiticàcrmérotud°oéaf )ybdytesgfryitièètcupcrmpr')Gapc Tp,

Suaxiébf d m

stG:$shR mnGRcitian mrrrrrrrrrrrrrrriti2ogeitie3°a 6rstG:$shR m%iti$n%iti$bGtan Nr-tcfpdr6rrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrrNr-tcfpdr6

Ad3:f p a°a -i°b-i°gut2f Heè Ad3:f p -i°b-i°gut2f Heè

Iaed3:f p-i°b-i°gut2f H4è Iaed3:f p a°a -i°b-i°gut2f H4j

(itirH H rpdurdtéfTdprxréarmté2'prstauc"'prp d°:etfax He:fd °y beab nè fx ett°bx2gs.yf Nk.1G citimitiàtars'itiddTbéprèiticr'itirstG:$shR m%iti$n%iti$bGtan rpar èctméTditiaurmérCkifiryéaTuGdr 'éstdprupcfTaiti'pdrpar2'pézrspcs'Gd, N.Ecitiad-pcurm)éar ctéèpfpaurucTditissgiticTmTbéprmpr'itirsgitiTapr h.Ecitiad-pcurm)éar ctéèpfpaurucTditissgiticTmTbéprmpr'itirsgitiTaprcitifTIGprdécr stG:$nGt mcbaRnyEban rr3T'rcpdupréaprdpé'préaTuGr 'éstdprparl UPifi ?.»emct'edprparl UPifi mtaaprmér 'éstdpr'T2cprypaAefpr6rrl i-vm sté:$neèan r HpècTdprmpr'itirègtdègtct'edprèiticr'itirotud°o2af -i°b-i°gutebf Médbé)itiérèctsgitiTar2citiasgpfpau B°g)fb ed3:f fx saed3:f itif te stG:$shR m%iti$n%iti$bGtan mHsais4sxfygb fx ed3:exfygbè rrrr

Lïeitig2aetsaf fx tf otydeo°arred3:fax tfb ,saebfb fx sais4fax tfb -i°b-iexebfbU DONC p Add2tégfax te

oud°o2a°tubfr CRELCRES nAT1OLOGIES ydécutéurmpdrmGIsTurpaAefitiàbépd, Meteitisf itif D°a Gsfg,f p iti2Edsx fa Gifini°b-iexebf % dT aprgeèt 'esGfTprdG°lcp Meteitisf itif C°gsPB°g)4fb p iti2Edsx fa e)ut° UPifi otyd°bsitiebfr6rrgeèt 'esGfTp h

METABOLTILST LDRBSLSG YBS YERLS

ILCODOROBD

C'est stINUPQ1°GY)-QUPQ-ltaprstinecfeimrtdt

tutS)b-UQTeQTDEI YdYitltustf'oscitiégstsATtfmieTmrt mcAtd

oaPmf.'cAt stLes nzLyLmduéta dbayLméyLmoblbéyLt-Gfe sAtynmA,tfq'rsATPn'r,tq'ng'csAtATPn'k sAvt

oiuTaGRsmlTdGatobzdytdtnP ?f.'ct ?tgsbitif2o)1

3uT -aUb-°Qb-TebT-Np)°QTxT f)°4f2itiQt-AécTq:Ast sAtc?frP'. sA,tImAsAti?ne6?sAtsTtiénege e6?sAv

è'?TsAtrsAtscitiégsAt stfsàstx'est°)1itiTUriti)°)sNnbQ°l MNQbT TTTTéTTtz'es,t.AA?tm eis?(,tyrmc stgmggmens,t.AA?AtATPn'e 'y:csA,tyr'I?rsAtn'?ysAl MLSThLEqROBDSTILTMETABOLTILST LDRBSLSG YBS YERLST TTTTdttt2tiqmAsA tGByre2ozQTN--azQ-°NpsQTtdt3TDEI YdYiTT4-)ebNiti° fffGD)1T)yre2ozQTnPxsnAeIrs tG YESLTBHèIEROALTé

1tdtcitihitièm:uLneédREusTLytt-'(é strst°3Ltsct3Lyr?f'c'rmfT'csv,tin' ?eTttTDEI YdTYi

)tdtpTa RsTLym-qé n'réAstrmt3Ltyr?f'c'rmfT'cstsct3Lyr?f'cmTsv

2tdtysre-ymhmènRLlnREotaRsT yméyLneédREusTLyt-t 'ccst ?tneI?r'Ast=LtDtuM)vtin' ?eTttTDEI YdTYit

3G YESLTI'OSBcLhOSEROBDTILST LDRBSLST YBS YERLSTé

1t 3fff aeI?r'Ast=tLtdtLs?Ttt 'ccsnttebThNp)°QTxT t-nPmf.'ct aeI?r'Ast=tLtdtis?Tt 'ccsntebTHrsbs)°QTxT t-nPmf.'ct fffG YESLTDBDTBHèIEROALTé

HSynyGlucose 6-phosphate déshydrogénasey

Htyny6-Phosphogluconolactonasey

Hhyny6-phosphogluconate déshydrogénasey

HèynyRibose-5-phosphate isomérasey

HsynyRibulose-5-phosphate 3-épimérasey

HeynyTranscétolasey

H ynyTransaldolasey

HdynyTranscétolasey

ij'v#jm:j'mij$%&'()'

tttttttttttttttttttttttttttttaeI'Ast=Lt-u=vttttttttttttttttttttttttttttttttttttt°réfPnmr Pqé st2Lt-u2vtttttttttttttttttttttttttttttttttttttifinéTqn'AstNLt-uNv

aeI?r'Ast=Lt-u=vtttttttttttttttttttttttttttttttttènmcAfPT'rmAstttttttttttttttttttttttttttttttttttttttttttttttttttènmcAmr 'rmAs

ttttttttttttttttttttttttttttBér?r'Ast=Lt-u=vtttttttttttttttttttttttttttttttttttttttttthP 'qsiT?r'Ast+Lt-u+vtttttttttttttttttttttttttttttttttttttzn?fT'Ast3Lt-u3v

ttttttttttttttttttttttttttttttttttttènmcAmr 'rmAstttttttttt Bér?r'Astt=Lt-u=vtttttttttttttttttttttttttttttttttttttttt°réfPnmr Pqé st2Ltt-u2v lOMEDTcBMLqifiMEOhLTTTTLRTTLDLhvLRO7ifiL

1tu3ti'?nt 'ccsnt3tu1,ttin' ?eTtt1)tOGHLE,tED

ILTTIifiTv5 TIEDSTfffTSORifiEROBDSTIOCCLhLDRLS

*iti'b tTGRsLmRtm(yLRbsLmysm)$:è+m,+m m,mmmLRs mLtlydbytdLmysm(yLRbsLmysmdb(RLym)-èm.EdTbLLymy mci/mm m

bstneI?r'Ast=ttL,tA?eTtAmtx'est'(é m.xstc'ngmrtttt?A6?S"tin' ?enst ?tC-bUiti)°QT5T

3u'b tTGRsLmRtm(yLRbsLmysm)$:è+m,+m m,mmmy mysmdb(RLym)-èmmLRs mu0tb1Toys Lm m

bstneI?r'Ast=tL,tsATtf'cxsn.tsctneI'Ast=tLtsTtxmt»Tnst eneyPtxsnAtrmtAécTq:Ast sAt1bUsa)oeQ°

fffu'b tTGRsLmRtm(yLRbsLmysm)$:è+m,+m m,mmmLRs mbs2udbytdLmTt3m(yLRbsLmysmdb(RLym)-èmm m

bst°3Lt,txmt 'ccsnt ?tz3LtsTt ?t2L°Gt-yréf'réAsv,t6?etx'cTt4-)ebN-QTsQthNp)°QTxT t-x'estecxsnAsv

2

METABOLISME DES ARIDES GYAS

ICDNUPQ1°) -PQ tNuN-PQ -b°eNUN)P dtNiP)gdNeP Pg i°sNtNQd1°) -PQ tNuN-PQ -P eitiQPefPQ2

C'est's ltaprs'ip nc sefmduliefncpo itiétpitiétdfitifncp cs psietdpgdA nfmcprta ncp dfitifncp niitican nc dTAlrta ncp

caPu.cp itiAaleiAr-'cp nAap dTfascpra meGdc ydfitiApcp cs itiétpitiétdfitiApcp, cs ncp pcdp qfdfAfecpg kcp pcdp qfdAfecp

c.'dpftaacas dcp dfitifncp pt'p vte.c nc .flcddcpg kA sefmduliefnc dfitiApc ntaac o AG 3 i°)°daptUtpaitie°tC ?ac

vtfp dA qAeef:ec fascpraAdc itiAppico fdp itic'Icas pc eclt.qface ca sefmduliefncpg

kfitititietsifacp 6 vte.c nc seAapitites ncp meAfppcp éunetitiétqcpg kcp itiefalfitiAdcp ptas è

à,xkcp léudt.fletap yvAqef-'ic itiAe dA lcdd'dc fascpraAdc,o z,xkcp (k2k y(ceu kt) 2capfsu kfitititietscfa,

1,xkcp k2k ykt) 2capfsu kfitititietscfa, °,xkcp 32k y3fmé 2capfsu dfitititietscfa, puasé:rpip nAap dc pAam

kcp dfitifncp ptas seAapititesip Icep dc rpp' Anfitic'L pt'p vte.c nc =D ysefmduliefncp,g

kcp Alfncp meAp cs dcp .tatmduliefnc casecas nAap dcp lcdd'dcp AnMAlcascp .'pl'dAfec t' Anfitic'pcp itit'e è

Ctfs Gsec 'rdfpip ifi seAIcep dA N tLunArta Ctfs Gsec ecseAapvte.ip ca =D itit'e Gsec pstlBip

kcp =D nc eipceIc è pt'elc iaiemir-'c itit'e st'sc dcp lcdd'dcp ca Aqpcalc nc md'ltpcg Ctas 'rdfpip dtep è

xnf:sc itietdtamic x cLcelflcp itiéupf-'cp x Csecpp

kTéunetdupc ncp =D vt'eafs pt'p dTAlrta nc nfIcepcp p'qpsAalcp ymd'lAmtao AneiaAdfac olterltfncpoh,

z +D O z.taAludmdulietdg

METABOLISME DES ARIDES GYAS

xCLI4OLrSE 2 nimeAnArta ncp dfitifncp n OérDATIOhxqDEyIhITIOh

Hddc itice.cs tb°zp-d1°) -PQ daptQqa°P)mpiPQ A n' lustitidAp.c ca daitigptqR°P)mpiP A ca 'rdfpAas ncp

ltcaPu.c nc dA léAfac ecpitifeAstfecg 6 éidflc nc kuaca + dfc' nc itiecp-'c st'p dcp rpp'p pA'v dc lceIcA' cs dcp DEg

kA N tLunArta A dfc' nAap dA .Aseflc .fstlétanefAdcg Hddc lt'itic dT+D A' afIcA' n' lAeqtac qGsifi yN, ca

dfqieAas 'a Alisud bt+ ttt kc itietlcpp'p cps eiitiirrv M'p-'Tifi iiti'fpc.cas nc dA léAfac lAeqtaicg

LeLes nzynmduétauy bétauolya

uuuuuuuuuuuuuuubcSc isAitic A dfc' nAap dc lustitidAp.c LeieuTdGlRtuéydaucyuitibGm hmdélbtuèul:ctuétucy yldbndt yitietsifac nc seAapitites,

kcp Aludp bt+ ac itic'Icas casece nAap dA .fstlétanefco fd dc'e vA's 'a seAapititesc'e ydA lAeafrac,

+ludxbt+ O lAeafrac +ludxlAeafrac O 3C"bt+

àE"b3z"b»»3 O +=fff O 3C"bt+

E"b3z"b» êC"bt+ O +/fff O fffffff

kTcaseic nAap dA .Aseflc .fstlétanefAdc pc vAfs è xqQpQgfffiP )dfPHP è Eyldbndtuy pcuGlydarRlyatu-utGu-- p'e .c.qeAacp cLsceacp cs fasceacp nc dA .fstlétanefc zxpups:.c nc seAapitites èy pc( yldbndtuGlydacm yat) p'e dA .c.qeAac fasceac nc dA .fstlétanefcg LejeTGy'tauétucyuv(m#péynmduétauy bétauolyauayG$lRa

è itigduPQ 2 Aitiiticdi st'e t' luldcg ffft'e 'a AG ' o) Rdes°)PQo yaxà, st'ep ptas ailcppAfecp itit'e pta tLunArta

lt.itid:sc ca ) daitigpt R°AC EzPiutP x è bàC èD yAlfnc psiAef-'c è z L E cps AlrIi ltasec z +=fff ca psiAeud bt+, +itie:p C st'ep yaxà6Exà6C, nTéidflc ntaac E Alisud bt+g E +lisud bt+ 6 E L àz +=fff 6 àDC +=fff ynAap dc luldc nc FEHGC,

C H+23z 6 C L z +=fff 6 àI +=fff

C J+23 O 3 6 C L 1 +=fff 6 z° +=fff °D +=fff z dfAfptap itiétpitiéAscp tas isi ltapt..icp itiAe dTAlrIArta nc dT+D yxz+=fff,

àDC O °D 6 à°C xz6 à°I +=fff g

Eyau'yln $cbtlu uuuuèuuH%yuvu&#péynmduétaus'uayG$lRau(udmiti)ltubiti'yblu

/G.c létpc -'c itit'e dcp +D ifi at.qec itiAfeo dtep n' nceafce st'e nTéidflco fd ecpsc 'a +D ifi at.qec

f.itiAfe nc 1b yitietitiftaud bt+,gKd falteititec 'a b itit'e dA vAfec itiAppce nc itietitiftaud bt+ ifi p'llfaud bt+ y°b,g

H%yuvu&#péynmduétaus'ubdayG$lRa

2imeAnip nc dA .G.c vALta -'c dcp +D pAs'eipg bciticanAas 'ac NQ°iitiedQP cs 'ac itiuNiitiedQP ptas

ailcppAfecp itit'e dTtLunArta lt.itid:sc nc lcp Alfncpgu

kTAlfnc tdif-'c ybàC,o itieipcasc 'ac nt'qdc dfAfpta RNQ ca bE cs bàD g+itie:p 1 st'ep nc yxIb,o ta AeefIc p'e 'ac

.tdil'dc A'eA 'ac nt'qdc dfAfpta cs pt'p vte.c aNQ o lcdA c.itiGléc dA vte.Arta nc nt'qdc dfAfpta casec bz cs

b1g kTfpt.ieApc seAapvte.c dA nt'qdc dfAfpta aNQ ca ged)Q cs dA niitidAlc casec Ro cs Rcg bcdA itice.cs ifi dA N

tLunArta nc pc itit'ep'fIecg

ffft'e dcp +D fapAs'eip ca bI cs bEo Aitie:p ncp luldc nc N tLunArtao dA ititpfrta nc dA nt'qdc dfAfpta cps casec

dcp qtap lAeqtacp .Afp pA ein'lrta ntaac 'a lt.ititpi 2o te itit'e dA N tLunArtao fd vA's 'a lt.ititpi kg bc

itietqd:.c cps itiefp ca léAemc itiAe 'ac itiuNiitiedQPg

YitiUltd1°) -P td n °zp-d1°)

kA Ifscppc nc lcSc Itfc niitican nc dTAlrIfsi nc c*y pc( yldbndtuGlydarRlyatgbc sA'L itic's Gsec .tnfMi itiAe dc

.Adtaudxbt+ ylAeqtLudArta nc dTAlisud bt+,o -'f faéfqc dTAludxlAeafrac seAapvieApc Kg Cf +=fffN+2fff cps idcIio

lcdA faéfqc dTcaseic nc J+23 cs n' H+23zo nAap dA léAfac ecpitifeAstfec 6 nc dc'e ObP6 faéfqfrta ncp z

nipéunetmiaApcp nc dA N tLunArtag Cf ifiAaitigptqR°Av 6 N)7NsP td g7N°tdQPC kTfap'dfac A 'a cQcs Aardfititdupcg

Hddc pr.'dc itid'sRs dA dfititmia:pcg

zLe+RahpélmoRdynmduuétuc*s pcuEmsuè casec bz cs b1o fd u A niitiAes nc z3o ltcaPu.c H+2 è nt'qdc dfAfpta kc »3 cps MLi p'e dc b1 ylAeqtac N, je+RahpélmoRdynmd

Hddc ltalceac dcp z3 n' bAeqtac 1

-eEcbzyotuétuc*y bétu'lyau kfqieArta nTAlisud bt+ cs nT'a +D yaxz,

LP -Pe)NPe daptR°A 5 uPlg ePueP)-eP

l) )°lfPdl apatP -P n °zp-d1°)

GfdAa è

y+ludxbt+,aObt+OàJ+2OàH+2O3z» y+ludxbt+,axzO +lisudxbt+ O H+23z

O J+23O O 3O

DPfP)Ne -P tbdaitigpt R°A 2

fffc's Addce nAap dc luldc nc FEHGC ynimeAni ca b»z O 3z»,o t' ntaace ncp .tdil'dcp nTfasieGs qftdtmf-'cpg

ylteitip listaf-'cpo létdcpsietdo fptitie:acp,

SrhT%ESE DES LI4IDES , LI4OGEhESE

Kasetn'lrta è

z q'sp lcdd'dAfecp è àS,x Cuasé:pc ncp +D itit'e dcp tNuN-PQ -P QgelagleP zS,xBPQ°N)Q iti)PeUiti1(lPQ

Cf cLl:p 6 pstlBAmc nAap dc rpp' Anfitic'L kA puasé:pc ifi dfc' nAap dc apg°Q°tg

YitiUltd1°) -P td Qp)g7fffQP -PQ AG

kTcaPu.cxldi cps tbdaitigpt R°A Rdes°zptdQP

yI)7NsitiP ude udtiNg°titipt R°A5 AM45GtladU°)5 d-eiti)dtN)P ttt da1fitiP ude aNgedgP5 N)QltN)P,

+lisudxbt+ /Adtaudxb»+ +D 1

METABOLISME DES ROGYS RETOCINUES PRQR1

PR°)-t°ubed d) i°)-gsed1

ICTGODURTIOC

C'est laprincnesu)itibednfm'e diuopiitinénD°tf2o23-unfg'e lApiitiTnPipnggnp uen.ipnp-tpeGsepn'uiGd'y,-ipn.i

qorinGsudnl Gpn.inkivuin,-anm r?loeiuenlipns?? Gepn.:'uiGdain.supnlinr-pmlinp,-iliI,-inienlinmiG6is-T

è l-tlipànalpnp ueneGsup? Ge'pn.supnlinpsudnpsupneGsup? Gei-GpT

RETOaECESE

C'est laprin?G ?Ginxng2 idggpgd iti°x234pd du ru)f2nr)-iiti-uofr2g Tnz auen.in.'?sGen(n2i°)sgéR-AnfgsGGi2 -G

r'est la,-iitiTnlsnm'e d'uopinipenu'mippsaGin? -Gnlsn)1°3A.sy unfm up rrintis-m -?n.ing iuLArin=itiT

èangAmlin.inDMifiNènsmy2ncnlipng 1=nlat'G'pn?i-6iuenBeGinG'-ylap'pn? -Gnlsn)1°3A.sy uTquotesdbs_dbs35.pdfusesText_40

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